贮运中马拉巴栗长期处于黑暗条件下，气孔开度减小或者关闭，也会引起叶片黄化、衰老。
P. macrocarpa during the storage and transportation is in a dark condition for a long time , and the stomatal aperture minishes or close totally , which may leads to the etiolation and senescence of leaves .
马栗