这是因为在厌氧条件下，菌体生长来源的ATP主要由发酵途径的底物水平磷酸化偶联生成。
Under anaerobic conditions , the ATP required for biomass generation was mainly supplied through substrate phosphorylation in fermentation pathways .
底物水平磷酸化底物水平磷酸化底物水平磷酸化