在此基础上，结合形态分析，提出了绝大多数栽培牡丹起源于多个野生原种的观点.矮牡丹、紫斑牡丹、杨山牡丹是最主要的起源种，卵叶牡丹的作用较小。
Based on this , combining with analysis with morphology , the authors put forward that most of the Chinese tree peony cultivars were origined from hybridization by many species .
杨山